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@ARTICLE{Lammich:136395,
author = {Lammich, Sven and Kamp, Frits and Wagner, Judith and
Nuscher, Brigitte and Zilow, Sonja and Ludwig, Ann-Katrin
and Willem, Michael and Haass, Christian},
title = {{T}ranslational repression of the disintegrin and
metalloprotease {ADAM}10 by a stable {G}-quadruplex
secondary structure in its 5'-untranslated region.},
journal = {The journal of biological chemistry},
volume = {286},
number = {52},
issn = {0021-9258},
address = {Bethesda, Md.},
publisher = {Soc.60645},
reportid = {DZNE-2020-02717},
pages = {45063-45072},
year = {2011},
abstract = {Anti-amyloidogenic processing of the amyloid precursor
protein APP by α-secretase prevents formation of the
amyloid-β peptide, which accumulates in senile plaques of
Alzheimer disease patients. α-Secretase belongs to the
family of a disintegrin and metalloproteases (ADAMs), and
ADAM10 is the primary candidate for this anti-amyloidogenic
activity. We recently demonstrated that ADAM10 translation
is repressed by its 5'-UTR and that in particular the first
half of ADAM10 5'-UTR is responsible for translational
repression. Here, we asked whether specific sequence motifs
exist in the ADAM10 5'-UTR that are able to form complex
secondary structures and thus potentially inhibit ADAM10
translation. Using circular dichroism spectroscopy, we
demonstrate that a G-rich region between nucleotides 66 and
94 of the ADAM10 5'-UTR forms a highly stable,
intramolecular, parallel G-quadruplex secondary structure
under physiological conditions. Mutation of guanines in this
sequence abrogates the formation of the G-quadruplex
structure. Although the G-quadruplex structure efficiently
inhibits translation of a luciferase reporter in in vitro
translation assays and in living cells, inhibition of
G-quadruplex formation fails to do so. Moreover, expression
of ADAM10 was similarly repressed by the G-quadruplex.
Mutation of the G-quadruplex motif results in a significant
increase of ADAM10 levels and consequently APPsα secretion.
Thus, we identified a critical RNA secondary structure
within the 5'-UTR, which contributes to the translational
repression of ADAM10.},
keywords = {5' Untranslated Regions: physiology / ADAM Proteins:
biosynthesis / ADAM Proteins: genetics / ADAM10 Protein /
Amyloid Precursor Protein Secretases: biosynthesis / Amyloid
Precursor Protein Secretases: genetics / HEK293 Cells /
Humans / Membrane Proteins: biosynthesis / Membrane
Proteins: genetics / Mutation / Nucleic Acid Conformation /
Protein Biosynthesis: physiology / 5' Untranslated Regions
(NLM Chemicals) / Membrane Proteins (NLM Chemicals) /
Amyloid Precursor Protein Secretases (NLM Chemicals) / ADAM
Proteins (NLM Chemicals) / ADAM10 Protein (NLM Chemicals) /
ADAM10 protein, human (NLM Chemicals)},
cin = {AG Haass old},
ddc = {540},
cid = {I:(DE-2719)1110007},
pnm = {342 - Disease Mechanisms and Model Systems (POF3-342)},
pid = {G:(DE-HGF)POF3-342},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:22065584},
pmc = {pmc:PMC3248004},
doi = {10.1074/jbc.M111.296921},
url = {https://pub.dzne.de/record/136395},
}