TY - JOUR AU - Jadavji, Nafisa M AU - Farr, Tracy D AU - Lips, Janet AU - Khalil, Ahmed A AU - Boehm-Sturm, Philipp AU - Foddis, Marco AU - Harms, Christoph AU - Füchtemeier, Martina AU - Dirnagl, Ulrich TI - Elevated levels of plasma homocysteine, deficiencies in dietary folic acid and uracil-DNA glycosylase impair learning in a mouse model of vascular cognitive impairment. JO - Behavioural brain research VL - 283 SN - 0166-4328 CY - Amsterdam PB - Elsevier M1 - DZNE-2020-04176 SP - 215-226 PY - 2015 AB - Dietary deficiencies in folic acid result in elevated levels of plasma homocysteine, which has been associated with the development of dementia and other neurodegenerative disorders. Previously, we have shown that elevated levels of plasma homocysteine in mice deficient for a DNA repair enzyme, uracil-DNA glycosylase (UNG), result in neurodegeneration. The goal of this study was to evaluate how deficiencies in folic acid and UNG along with elevated levels of homocysteine affect vascular cognitive impairment, via chronic hypoperfursion in an animal model. Ung(+/+) and Ung(-/-) mice were placed on either control (CD) or folic acid deficient (FADD) diets. Six weeks later, the mice either underwent implantation of microcoils around both common carotid arteries. Post-operatively, behavioral tests began at 3-weeks, angiography was measured after 5-weeks using MRI to assess vasculature and at completion of study plasma and brain tissue was collected for analysis. Learning impairments in the Morris water maze (MWM) were observed only in hypoperfused Ung(-/-) FADD mice and these mice had significantly higher plasma homocysteine concentrations. Interestingly, Ung(+/+) FADD produced significant remodeling of the basilar artery and arterial vasculature. Increased expression of GFAP was observed in the dentate gyrus of Ung(-/-) hypoperfused and FADD sham mice. Chronic hypoperfusion resulted in increased cortical MMP-9 protein levels of FADD hypoperfused mice regardless of genotypes. These results suggest that elevated levels of homocysteine only, as a result of dietary folic acid deficiency, don't lead to memory impairments and neurobiochemical changes. Rather a combination of either chronic hypoperfusion or UNG deficiency is required. KW - Animals KW - Basilar Artery: pathology KW - Basilar Artery: physiopathology KW - Brain: blood supply KW - Brain: pathology KW - Brain: physiopathology KW - Carotid Artery Diseases KW - Cerebrovascular Disorders: pathology KW - Cerebrovascular Disorders: physiopathology KW - Chronic Disease KW - Cognition Disorders: pathology KW - Cognition Disorders: physiopathology KW - Diet KW - Disease Models, Animal KW - Female KW - Folic Acid Deficiency: pathology KW - Folic Acid Deficiency: physiopathology KW - Glial Fibrillary Acidic Protein KW - Gliosis: pathology KW - Gliosis: physiopathology KW - Homocysteine: blood KW - Learning Disabilities: pathology KW - Learning Disabilities: physiopathology KW - Male KW - Matrix Metalloproteinase 9: metabolism KW - Maze Learning: physiology KW - Mice, Inbred C57BL KW - Mice, Knockout KW - Nerve Tissue Proteins: metabolism KW - Random Allocation KW - Uracil-DNA Glycosidase: deficiency KW - Uracil-DNA Glycosidase: genetics KW - Glial Fibrillary Acidic Protein (NLM Chemicals) KW - Nerve Tissue Proteins (NLM Chemicals) KW - glial fibrillary astrocytic protein, mouse (NLM Chemicals) KW - Homocysteine (NLM Chemicals) KW - Uracil-DNA Glycosidase (NLM Chemicals) KW - Matrix Metalloproteinase 9 (NLM Chemicals) KW - Mmp9 protein, mouse (NLM Chemicals) LB - PUB:(DE-HGF)16 C6 - pmid:25655513 DO - DOI:10.1016/j.bbr.2015.01.040 UR - https://pub.dzne.de/record/137854 ER -