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@ARTICLE{Woeste:140584,
author = {Woeste, Marina A and Stern, Sina and Raju, Diana N and
Grahn, Elena and Dittmann, Dominik and Gutbrod, Katharina
and Dörmann, Peter and Hansen, Jan N and Schonauer, Sophie
and Marx, Carina E and Hamzeh, Hussein and Körschen, Heinz
G and Aerts, Johannes M F G and Bönigk, Wolfgang and
Endepols, Heike and Sandhoff, Roger and Geyer, Matthias and
Berger, Thomas K and Bradke, Frank and Wachten, Dagmar},
title = {{S}pecies-specific differences in nonlysosomal
glucosylceramidase {GBA}2 function underlie locomotor
dysfunction arising from loss-of-function mutations.},
journal = {The journal of biological chemistry},
volume = {294},
number = {11},
issn = {0021-9258},
address = {Bethesda, Md.},
publisher = {Soc.60645},
reportid = {DZNE-2020-06906},
pages = {3853-3871},
year = {2019},
abstract = {The nonlysosomal glucosylceramidase β2 (GBA2) catalyzes
the hydrolysis of glucosylceramide to glucose and ceramide.
Mutations in the human GBA2 gene have been associated with
hereditary spastic paraplegia (HSP), autosomal-recessive
cerebellar ataxia (ARCA), and the Marinesco-Sjögren-like
syndrome. However, the underlying molecular mechanisms are
ill-defined. Here, using biochemistry, immunohistochemistry,
structural modeling, and mouse genetics, we demonstrate that
all but one of the spastic gait locus #46 (SPG46)-connected
mutations cause a loss of GBA2 activity. We demonstrate that
GBA2 proteins form oligomeric complexes and that
protein-protein interactions are perturbed by some of these
mutations. To study the pathogenesis of GBA2-related HSP and
ARCA in vivo, we investigated GBA2-KO mice as a mammalian
model system. However, these mice exhibited a high
phenotypic variance and did not fully resemble the human
phenotype, suggesting that mouse and human GBA2 differ in
function. Whereas some GBA2-KO mice displayed a strong
locomotor defect, others displayed only mild alterations of
the gait pattern and no signs of cerebellar defects. On a
cellular level, inhibition of GBA2 activity in isolated
cerebellar neurons dramatically affected F-actin dynamics
and reduced neurite outgrowth, which has been associated
with the development of neurological disorders. Our results
shed light on the molecular mechanism underlying the
pathogenesis of GBA2-related HSP and ARCA and reveal
species-specific differences in GBA2 function in vivo.},
keywords = {Glucosylceramidase / Animals / Biocatalysis / Cerebellar
Ataxia: genetics / Cerebellar Ataxia: metabolism / Humans /
Locomotion: genetics / Loss of Function Mutation / Mice /
Mice, Knockout / Spastic Paraplegia, Hereditary: genetics /
Spastic Paraplegia, Hereditary: metabolism / Species
Specificity / beta-Glucosidase: antagonists $\&$ inhibitors
/ beta-Glucosidase: deficiency / beta-Glucosidase: genetics
/ beta-Glucosidase: metabolism / beta-Glucosidase (NLM
Chemicals) / beta-glucosidase 2, mouse (NLM Chemicals) /
GBA2 protein, human (NLM Chemicals)},
cin = {AG Bradke},
ddc = {540},
cid = {I:(DE-2719)1013002},
pnm = {341 - Molecular Signaling (POF3-341)},
pid = {G:(DE-HGF)POF3-341},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:30662006},
pmc = {pmc:PMC6422076},
doi = {10.1074/jbc.RA118.006311},
url = {https://pub.dzne.de/record/140584},
}