TY - JOUR AU - Khan, Nida S AU - Lukason, Daniel P AU - Feliu, Marianela AU - Ward, Rebecca A AU - Lord, Allison K AU - Reedy, Jennifer L AU - Ramirez-Ortiz, Zaida G AU - Tam, Jenny M AU - Kasperkovitz, Pia V AU - Negoro, Paige E AU - Vyas, Tammy D AU - Xu, Shuying AU - Brinkmann, Melanie M AU - Acharaya, Mridu AU - Artavanis-Tsakonas, Katerina AU - Frickel, Eva-Maria AU - Becker, Christine E AU - Dagher, Zeina AU - Kim, You-Me AU - Latz, Eicke AU - Ploegh, Hidde L AU - Mansour, Michael K AU - Miranti, Cindy K AU - Levitz, Stuart M AU - Vyas, Jatin M TI - CD82 controls CpG-dependent TLR9 signaling. JO - The FASEB journal VL - 33 IS - 11 SN - 0892-6638 CY - Hoboken, NJ PB - Wiley M1 - DZNE-2020-00423 SP - 12500-12514 PY - 2019 AB - The tetraspanin CD82 is a potent suppressor of tumor metastasis and regulates several processes including signal transduction, cell adhesion, motility, and aggregation. However, the mechanisms by which CD82 participates in innate immunity are unknown. We report that CD82 is a key regulator of TLR9 trafficking and signaling. TLR9 recognizes unmethylated cytosine-phosphate-guanine (CpG) motifs present in viral, bacterial, and fungal DNA. We demonstrate that TLR9 and CD82 associate in macrophages, which occurs in the endoplasmic reticulum (ER) and post-ER. Moreover, CD82 is essential for TLR9-dependent myddosome formation in response to CpG stimulation. Finally, CD82 modulates TLR9-dependent NF-κB nuclear translocation, which is critical for inflammatory cytokine production. To our knowledge, this is the first time a tetraspanin has been implicated as a key regulator of TLR signaling. Collectively, our study demonstrates that CD82 is a specific regulator of TLR9 signaling, which may be critical in cancer immunotherapy approaches and coordinating the innate immune response to pathogens. KW - Active Transport, Cell Nucleus: drug effects KW - Active Transport, Cell Nucleus: genetics KW - Active Transport, Cell Nucleus: immunology KW - Animals KW - Cell Nucleus: genetics KW - Cell Nucleus: immunology KW - Cytokines: genetics KW - Cytokines: immunology KW - Endoplasmic Reticulum: genetics KW - Endoplasmic Reticulum: immunology KW - Endoplasmic Reticulum: pathology KW - Inflammation: genetics KW - Inflammation: immunology KW - Inflammation: pathology KW - Kangai-1 Protein: genetics KW - Kangai-1 Protein: immunology KW - Macrophages: immunology KW - Macrophages: pathology KW - Mice KW - Mice, Knockout KW - NF-kappa B: genetics KW - NF-kappa B: immunology KW - Oligodeoxyribonucleotides: pharmacology KW - RAW 264.7 Cells KW - Signal Transduction: drug effects KW - Signal Transduction: genetics KW - Signal Transduction: immunology KW - Toll-Like Receptor 9: genetics KW - Toll-Like Receptor 9: immunology KW - CPG-oligonucleotide (NLM Chemicals) KW - Cd82 antigen, mouse (NLM Chemicals) KW - Cytokines (NLM Chemicals) KW - Kangai-1 Protein (NLM Chemicals) KW - NF-kappa B (NLM Chemicals) KW - Oligodeoxyribonucleotides (NLM Chemicals) KW - Tlr9 protein, mouse (NLM Chemicals) KW - Toll-Like Receptor 9 (NLM Chemicals) LB - PUB:(DE-HGF)16 C6 - pmid:31408613 C2 - pmc:PMC6988855 DO - DOI:10.1096/fj.201901547R UR - https://pub.dzne.de/record/145064 ER -