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@ARTICLE{Hochmair:163787,
author = {Hochmair, Janine and Exner, Christian and Franck,
Maximilian and Dominguez-Baquero, Alvaro and Diez, Lisa and
Brognaro, Hévila and Kraushar, Matthew L and Mielke,
Thorsten and Radbruch, Helena and Kaniyappan,
Senthilvelrajan and Falke, Sven and Mandelkow, Eckhard and
Betzel, Christian and Wegmann, Susanne},
title = {{M}olecular crowding and {RNA} synergize to promote phase
separation, microtubule interaction, and seeding of {T}au
condensates},
journal = {The EMBO journal},
volume = {41},
number = {11},
issn = {0261-4189},
address = {Hoboken, NJ [u.a.]},
publisher = {Wiley},
reportid = {DZNE-2022-00525},
pages = {e108882},
year = {2022},
note = {(CC BY-NC-ND)},
abstract = {Biomolecular condensation of the neuronal
microtubule-associated protein Tau (MAPT) can be induced by
coacervation with polyanions like RNA, or by molecular
crowding. Tau condensates have been linked to both
functional microtubule binding and pathological aggregation
in neurodegenerative diseases. We find that molecular
crowding and coacervation with RNA, two conditions likely
coexisting in the cytosol, synergize to enable Tau
condensation at physiological buffer conditions and to
produce condensates with a strong affinity to charged
surfaces. During condensate-mediated microtubule
polymerization, their synergy enhances bundling and spatial
arrangement of microtubules. We further show that different
Tau condensates efficiently induce pathological Tau
aggregates in cells, including accumulations at the nuclear
envelope that correlate with nucleocytoplasmic transport
deficits. Fluorescent lifetime imaging reveals different
molecular packing densities of Tau in cellular accumulations
and a condensate-like density for nuclear-envelope Tau.
These findings suggest that a complex interplay between
interaction partners, post-translational modifications, and
molecular crowding regulates the formation and function of
Tau condensates. Conditions leading to prolonged existence
of Tau condensates may induce the formation of
seeding-competent Tau and lead to distinct cellular Tau
accumulations.},
keywords = {Humans / Microtubules: metabolism / Neurodegenerative
Diseases: metabolism / Neurons: metabolism / Protein Binding
/ RNA: metabolism / tau Proteins: metabolism},
cin = {AG Wegmann / AG Mandelkow 1 / AG Mandelkow 2},
ddc = {570},
cid = {I:(DE-2719)1810006 / I:(DE-2719)1013014 /
I:(DE-2719)1013015},
pnm = {352 - Disease Mechanisms (POF4-352)},
pid = {G:(DE-HGF)POF4-352},
typ = {PUB:(DE-HGF)16},
pmc = {pmc:PMC9156969},
pubmed = {pmid:35298090},
doi = {10.15252/embj.2021108882},
url = {https://pub.dzne.de/record/163787},
}