TY - JOUR AU - Henning, Lukas AU - Antony, Henrike AU - Breuer, Annika AU - Müller, Julia AU - Seifert, Gerald AU - Audinat, Etienne AU - Singh, Parmveer AU - Brosseron, Frederic AU - Heneka, Michael T. AU - Steinhäuser, Christian AU - Bedner, Peter TI - Reactive microglia are the major source of tumor necrosis factor alpha and contribute to astrocyte dysfunction and acute seizures in experimental temporal lobe epilepsy JO - Glia VL - 71 IS - 2 SN - 0894-1491 CY - Bognor Regis [u.a.] PB - Wiley-Liss M1 - DZNE-2022-01566 SP - 168 - 186 PY - 2023 N1 - CC BY-NC-ND: https://creativecommons.org/licenses/by-nc-nd/4.0/ AB - Extensive microglia reactivity has been well described in human and experimental temporal lobe epilepsy (TLE). To date, however, it is not clear whether and based on which molecular mechanisms microglia contribute to the development and progression of focal epilepsy. Astroglial gap junction coupled networks play an important role in regulating neuronal activity and loss of interastrocytic coupling causally contributes to TLE. Here, we show in the unilateral intracortical kainate (KA) mouse model of TLE that reactive microglia are primary producers of tumor necrosis factor (TNF)α and contribute to astrocyte dysfunction and severity of status epilepticus (SE). Immunohistochemical analyses revealed pronounced and persistent microglia reactivity, which already started 4 h after KA-induced SE. Partial depletion of microglia using a colony stimulating factor 1 receptor inhibitor prevented early astrocyte uncoupling and attenuated the severity of SE, but increased the mortality of epileptic mice following surgery. Using microglia-specific inducible TNFα knockout mice we identified microglia as the major source of TNFα during early epileptogenesis. Importantly, microglia-specific TNFα knockout prevented SE-induced gap junction uncoupling in astrocytes. Continuous telemetric EEG recordings revealed that during the first 4 weeks after SE induction, microglial TNFα did not significantly contribute to spontaneous generalized seizure activity. Moreover, the absence of microglial TNFα did not affect the development of hippocampal sclerosis but attenuated gliosis. Taken together, these data implicate reactive microglia in astrocyte dysfunction and network hyperexcitability after an epileptogenic insult. KW - Mice KW - Animals KW - Humans KW - Epilepsy, Temporal Lobe: pathology KW - Astrocytes: pathology KW - Tumor Necrosis Factor-alpha KW - Microglia: pathology KW - Hippocampus: pathology KW - Seizures: pathology KW - Status Epilepticus: pathology KW - Kainic Acid: toxicity KW - Disease Models, Animal KW - Mice, Knockout KW - astrocyte (Other) KW - gap junction coupling (Other) KW - hippocampal sclerosis (Other) KW - microglia (Other) KW - temporal lobe epilepsy (Other) KW - tumor necrosis factor alpha (Other) KW - Tumor Necrosis Factor-alpha (NLM Chemicals) KW - Kainic Acid (NLM Chemicals) LB - PUB:(DE-HGF)16 C6 - pmid:36373840 DO - DOI:10.1002/glia.24265 UR - https://pub.dzne.de/record/165273 ER -