TY - JOUR AU - Dawkins, Edgar AU - Derks, Rico J E AU - Schifferer, Martina AU - Trambauer, Johannes AU - Winkler, Edith AU - Simons, Mikael AU - Paquet, Dominik AU - Giera, Martin AU - Kamp, Frits AU - Steiner, Harald TI - Membrane lipid remodeling modulates γ-secretase processivity. JO - The journal of biological chemistry VL - 299 IS - 4 SN - 0021-9258 CY - Bethesda, Md. PB - Soc. M1 - DZNE-2023-00500 SP - 103027 PY - 2023 AB - Imbalances in the amounts of amyloid-β peptides (Aβ) generated by the membrane proteases β- and γ-secretase are considered as a trigger of Alzheimer's disease (AD). Cell-free studies of γ-secretase have shown that increasing membrane thickness modulates Aβ generation but it has remained unclear if these effects are translatable to cells. Here we show that the very long-chain fatty acid erucic acid (EA) triggers acyl chain remodeling in AD cell models, resulting in substantial lipidome alterations which included increased esterification of EA in membrane lipids. Membrane remodeling enhanced γ-secretase processivity, resulting in the increased production of the potentially beneficial Aβ37 and/or Aβ38 species in multiple cell lines. Unexpectedly, we found that the membrane remodeling stimulated total Aβ secretion by cells expressing WT γ-secretase but lowered it for cells expressing an aggressive familial AD mutant γ-secretase. We conclude that EA-mediated modulation of membrane composition is accompanied by complex lipid homeostatic changes that can impact amyloidogenic processing in different ways and elicit distinct γ-secretase responses, providing critical implications for lipid-based AD treatment strategies. KW - Humans KW - Amyloid Precursor Protein Secretases: genetics KW - Amyloid Precursor Protein Secretases: metabolism KW - Membrane Lipids: metabolism KW - Amyloid beta-Peptides: metabolism KW - Alzheimer Disease: genetics KW - Alzheimer Disease: metabolism KW - Cell Line KW - Amyloid beta-Protein Precursor: metabolism KW - Presenilin-1: metabolism KW - Aβ37/38 (Other) KW - Alzheimer disease (Other) KW - Aβ37/38 (Other) KW - amyloid precursor protein (APP) processing (Other) KW - amyloid-β peptide (Aβ) (Other) KW - erucic acid (Other) KW - lipid homeostasis (Other) KW - lipidomics (Other) KW - membrane thickness (Other) KW - presenilin (Other) KW - γ-secretase (Other) KW - Amyloid Precursor Protein Secretases (NLM Chemicals) KW - Membrane Lipids (NLM Chemicals) KW - Amyloid beta-Peptides (NLM Chemicals) KW - Amyloid beta-Protein Precursor (NLM Chemicals) KW - Presenilin-1 (NLM Chemicals) KW - amyloid-β peptide (Aβ) (Other) KW - γ-secretase (Other) LB - PUB:(DE-HGF)16 C6 - pmid:36805335 C2 - pmc:PMC10070668 DO - DOI:10.1016/j.jbc.2023.103027 UR - https://pub.dzne.de/record/257785 ER -