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@ARTICLE{Dunot:271713,
author = {Dunot, Jade and Moreno, Sebastien and Gandin, Carine and
Pousinha, Paula A and Amici, Mascia and Dupuis, Julien and
Anisimova, Margarita and Winschel, Alex and Uriot, Magalie
and Petshow, Samuel J and Mensch, Maria and Bethus, Ingrid
and Giudici, Camilla and Hampel, Heike and Wefers, Benedikt
and Wurst, Wolfgang and Naumann, Ronald and Ashby, Michael C
and Laube, Bodo and Zito, Karen and Mellor, Jack R and Groc,
Laurent and Willem, Michael and Marie, Hélène},
title = {{APP} fragment controls both ionotropic and non-ionotropic
signaling of {NMDA} receptors.},
journal = {Neuron},
volume = {112},
number = {16},
issn = {0896-6273},
address = {New York, NY},
publisher = {Elsevier},
reportid = {DZNE-2024-01065},
pages = {2708 - 2720.e9},
year = {2024},
abstract = {NMDA receptors (NMDARs) are ionotropic receptors crucial
for brain information processing. Yet, evidence also
supports an ion-flux-independent signaling mode mediating
synaptic long-term depression (LTD) and spine shrinkage.
Here, we identify AETA (Aη), an amyloid-β precursor
protein (APP) cleavage product, as an NMDAR modulator with
the unique dual regulatory capacity to impact both signaling
modes. AETA inhibits ionotropic NMDAR activity by competing
with the co-agonist and induces an intracellular
conformational modification of GluN1 subunits. This favors
non-ionotropic NMDAR signaling leading to enhanced LTD and
favors spine shrinkage. Endogenously, AETA production is
increased by in vivo chemogenetically induced neuronal
activity. Genetic deletion of AETA production alters NMDAR
transmission and prevents LTD, phenotypes rescued by acute
exogenous AETA application. This genetic deletion also
impairs contextual fear memory. Our findings demonstrate
AETA-dependent NMDAR activation (ADNA), characterizing AETA
as a unique type of endogenous NMDAR modulator that exerts
bidirectional control over NMDAR signaling and associated
information processing.},
keywords = {Humans / Receptors, N-Methyl-D-Aspartate: metabolism /
Animals / Mice / Amyloid beta-Protein Precursor: genetics /
Amyloid beta-Protein Precursor: metabolism / Signal
Transduction: physiology / Long-Term Synaptic Depression:
physiology / Long-Term Synaptic Depression: drug effects /
Mice, Inbred C57BL / Mice, Knockout / Fear: physiology /
Hippocampus: metabolism / Neurons: metabolism / Dendritic
Spines: metabolism / Memory: physiology / Rats / APP (Other)
/ NMDA receptors (Other) / amyloid-β precursor protein
(Other) / eta-secretase (Other) / hippocampus (Other) /
long-term depression (Other) / memory (Other) /
non-ionotropic signaling (Other) / spine shrinkage (Other) /
synapse (Other) / Receptors, N-Methyl-D-Aspartate (NLM
Chemicals) / Amyloid beta-Protein Precursor (NLM Chemicals)},
cin = {AG Haass / AG Wurst},
ddc = {610},
cid = {I:(DE-2719)1110007 / I:(DE-2719)1140001},
pnm = {352 - Disease Mechanisms (POF4-352)},
pid = {G:(DE-HGF)POF4-352},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:38878768},
pmc = {pmc:PMC11343662},
doi = {10.1016/j.neuron.2024.05.027},
url = {https://pub.dzne.de/record/271713},
}