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@ARTICLE{Ziegler:281537,
author = {Ziegler, Anna B and Wesselmann, Cedrik and Beckschäfer,
Konstantin and Wulf, Anna-Lena and Dhiman, Neena and Soba,
Peter and Thiele, Christoph and Bauer, Reinhard and
Tavosanis, Gaia},
title = {{I}ndividual lipid alterations at the origin of neuronal
{C}eramide {S}ynthase defects.},
journal = {PLoS Genetics},
volume = {21},
number = {9},
issn = {1553-7390},
address = {San Francisco, Calif.},
publisher = {Public Library of Science},
reportid = {DZNE-2025-01155},
pages = {e1011880},
year = {2025},
abstract = {The brain is highly susceptible to disturbances in lipid
metabolism. Among the rare, genetically-linked epilepsies
Progressive Myoclonic Epilepsy Type 8 (PME8), associated
with the loss of Ceramide Synthase (CerS) activity, causes
epileptic symptoms accompanied by early onset of
neurodegenerative traits. The function of CerS is embedded
in a complex, conserved metabolic pathway, making it
difficult to identify the specific disease-relevant
alterations. Here, we show that the expression of an
enzymatically inactive cerS allele in Drosophila sensory
neurons yielded developmental and early onset dendrite loss.
Combining lipidomics and refined genetics with quantitative
analysis of neuronal morphology in cerS mutants, we
identified which lipids species are dysregulated and how
they affect neuronal morphology. In cerS mutants, long and
very-long acyl-chain C18-C24-ceramides were missing and
necessary for dendrite elaboration. In addition, the
substrate of CerS, (dh)S, and its metabolite (dh)S1P,
increased. Especially increasing (dh)S1P strongly reduces
dendritic complexity in cerS mutant neurons. Finally, we
performed in vivo experiments to cell-autonomously rescue
the morphological defects of cerS mutant neurons and report
that a complete rescue can only be achieved if the toxic
CerS substrate is converted to produce specific (C18-C24)
ceramides. Thus, despite the complex metabolic alterations,
our data provides essential information about the metabolic
origin of PME8 and delineates a potential therapeutic
avenue.},
keywords = {Animals / Lipid Metabolism: genetics / Ceramides:
metabolism / Oxidoreductases: genetics / Oxidoreductases:
metabolism / Drosophila melanogaster: genetics / Dendrites:
metabolism / Dendrites: pathology / Dendrites: genetics /
Drosophila Proteins: genetics / Drosophila Proteins:
metabolism / Neurons: metabolism / Mutation / Lipidomics /
Drosophila / dihydroceramide desaturase (NLM Chemicals) /
Ceramides (NLM Chemicals) / Oxidoreductases (NLM Chemicals)
/ Drosophila Proteins (NLM Chemicals)},
cin = {AG Tavosanis},
ddc = {610},
cid = {I:(DE-2719)1013018},
pnm = {351 - Brain Function (POF4-351)},
pid = {G:(DE-HGF)POF4-351},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:40997116},
doi = {10.1371/journal.pgen.1011880},
url = {https://pub.dzne.de/record/281537},
}