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@ARTICLE{Gunkel:137875,
author = {Gunkel, Monika and Schöneberg, Johannes and Alkhaldi,
Weaam and Irsen, Stephan and Noé, Frank and Kaupp, U
Benjamin and Alamoudi, Ashraf},
title = {{H}igher-order architecture of rhodopsin in intact
photoreceptors and its implication for phototransduction
kinetics.},
journal = {Structure},
volume = {23},
number = {4},
issn = {0969-2126},
address = {Cambridge, Mass.},
publisher = {Cell Press},
reportid = {DZNE-2020-04197},
pages = {628-638},
year = {2015},
abstract = {The visual pigment rhodopsin belongs to the family of G
protein-coupled receptors that can form higher oligomers. It
is controversial whether rhodopsin forms oligomers and
whether oligomers are functionally relevant. Here, we study
rhodopsin organization in cryosections of dark-adapted mouse
rod photoreceptors by cryoelectron tomography. We identify
four hierarchical levels of organization. Rhodopsin forms
dimers; at least ten dimers form a row. Rows form pairs
(tracks) that are aligned parallel to the disk incisures.
Particle-based simulation shows that the combination of
tracks with fast precomplex formation, i.e. rapid
association and dissociation between inactive rhodopsin and
the G protein transducin, leads to kinetic trapping:
rhodopsin first activates transducin from its own track,
whereas recruitment of transducin from other tracks proceeds
more slowly. The trap mechanism could produce uniform
single-photon responses independent of rhodopsin lifetime.
In general, tracks might provide a platform that coordinates
the spatiotemporal interaction of signaling molecules.},
keywords = {Animals / Kinetics / Mice / Mice, Inbred C57BL /
Photoreceptor Cells: metabolism / Photoreceptor Cells:
ultrastructure / Protein Binding / Protein Multimerization /
Rhodopsin: chemistry / Rhodopsin: metabolism / Transducin:
metabolism / Vision, Ocular / Rhodopsin (NLM Chemicals) /
Transducin (NLM Chemicals)},
cin = {AG Alamoudi},
ddc = {540},
cid = {I:(DE-2719)1013012},
pnm = {341 - Molecular Signaling (POF3-341)},
pid = {G:(DE-HGF)POF3-341},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:25728926},
doi = {10.1016/j.str.2015.01.015},
url = {https://pub.dzne.de/record/137875},
}